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The fly is Tipula confusa, caught 7 October 2022 at Moelyci Farm, Tregarth, North Wales. It was quite lively when I caught it, but under the microscope I saw the tip of the abdomen was missing. In the cavity were some black spheres attached by thin strands - photo. I'm wondering if these are fungi and would they be the cause of the Tipula losing its abdomen tip?

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I would say these black spheres are spermathecae - the structures in which the female stores sperm before fertilization of the eggs. Quite why the fly should be missing its abdomen tip, I don't know.

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Thanks very much for the explanation.
John